Peer-reviewed publications striped mouse project

(Full publication list C. Schradin including publications from other projects)

70. Schradin C. 2017. Comparative studies need to rely both on sound natural history data and on excellent statistical analysis. Royal Society Open Science. 4: 170346.


69. Schradin, C. & Hayes, L.D. 2017. A synopsis of long-term field studies in mammals: achievements, future directions, and some advice Journal of Mammalogy, 98, 670-677.


68- Hayes, L.D., Ebensperger, L.A., Kelt, D.A., Meserve, P.L., Pillay, N., Viblanc, V.A. & Schradin, C. 2017. Long-term field studies in rodents. Journal of Mammalogy, 98, 642-651.


67. Hayes, L. D., & Schradin, C. 2017. Long-term field studies in mammals: what the short-term study cannot tell us. Journal of Mammalogy, 98, 600-602.


66. Jäger, J., C. Schradin, N. Pillay, and R. Rimbach. 2017. Active and explorative individuals are often restless and excluded from studies measuring resting metabolic rate: Do alternative metabolic rate measures offer a solution? Physiology & Behavior, 174,57-66.


65. Maille, A. & Schradin, C. 2017. Eco-physiology of cognition. Biological Reviews, 92, 1101-1112.


64. Schoepf, I., N. Pillay, and C. Schradin. 2017. Trade-offs between reproduction and health in free-ranging African striped mice. Journal of Comparative Physiology B 187:625-637.


63. Rimbach, R., M. Wastavino, C. H. Yuen, N. Pillay, and C. Schradin. 2017. Contrasting activity budgets of alternative reproductive tactics in male striped mice. Journal of Zoology 301:280-289.


62. Schoepf, I., N. Pillay, and C. Schradin. 2017. The pathophysiology of survival in harsh environments. Journal of Comparative Physiology B, 187,183-201.


61. Rimbach, R., N. Pillay, and C. Schradin. 2017. Both thyroid hormone levels and resting metabolic rate decrease in African striped mice when food availability decreases. The Journal of Experimental Biology.


60. Mallarino, R., Henegar, C., Mirasierra, Manceau, M., Schradin, C., Vallejo. M., Beronja, S., Barsh, G.S., Hoekstra, H.E: accept. Alx3 regulates the spatial differences in hair pigment underlying stripe patterns in rodents. Nature, 539, 518-523.


59 . Maille, A. & Schradin, C. 2016. Survival is linked with reaction time and spatial memory in African striped mice. Biology Letters 12, 10.1098/rsbl.2016.0346.


58. Rimbach, R., R. Willigenburg, I. Schoepf, C. H. Yuen, N. Pillay, and C. Schradin. 2016. Young But Not Old Adult African Striped Mice Reduce Their Activity in the Dry Season When Food Availability is Low. Ethology 122, 828-840.


57. Nater, C.R., Canale, C.I., van Benthem, K.J., Yuen, C.H., Schoepf, I., Pillay, N. Ozgul, A. & Schradin, C. (2016). Interactive effects of exogenous and endogenous factors on demographic rates of an African rodent. Oikos, 125, 1838-1848.


 56.Rymer, T. L., N. Pillay, and C. Schradin. 2016. Resilience to droughts in mammals: a conceptual framework for estimating vulnerability of a single species. The Quarterly Reviews of Biology 91, 133-176.


55. Yuen, C. H., N. Pillay, M. Heinrichs, I. Schoepf, and C. Schradin. 2016. Personality traits are consistent when measured in the field and in the laboratory in African striped mice (Rhabdomys pumilio). Behavioral Ecology and Sociobiology 70:1235-1246


54. Schoepf, I., Brunner, P., Yuen, C.H., König. B. & Schradin, C. 2016. Does a mouse have a friend? Mixed evidence for individual recognitio n in the African striped mouse (Rhabdomys pumilio). Journal of Zoology 299, 142-149.



53. Hill, D., Pillay, N. & Schradin, C. 2015. Alternative reproductive tactics in female striped mice: solitary breeders have lower corticosterone levels than communal breeders. Hormones and Behavior 71, 1-9.


52. Hill, D., Pillay, N. & Schradin, C. 2015. Alternative reproductive tactics in female striped mice: heavier females are more likely to breed solitary than communally. Journal of Animal Ecology 84, 1497-1508.


51. Maille, A., Pillay, N. & Schradin, C. 2015. Seasonal variation in attention and spatial performance in a wild population of the African striped mouse Rhabdomys pumilio. Animal Cognition 18, 1231-1242.


50. Raynaud, J. & Schradin, C. 2015. Corticosterone levels correlate with alloparental care in a sex-dependent manner in African striped mice, Rhabdomys pumilio. Ethology 121, 57-67.


49. Schoepf, I., Kenkel, W. & Schradin, C. 2015. Arginine vasopressin in brains of free ranging striped mouse males following alternative reproductive tactics, Journal of Ethology 33, 235-242.


48. Schoepf, I. Schmohl, G., König, B., Pillay, N., Schradin, C. 2015. Manipulation of population density and food availability affects home-range sizes of African striped mouse females. Animal Behavior 99, 53-60.


47. Schradin, C., Pillay, N., Kondratyeva, A., Chi-Hang, Y., Schoepf, I., Krackow, S. 2015. Basal blood glucose concentration in free living striped mice is influenced by food availability, ambient temperature and social tactic. Biology Letters.


46. Yuen, C.H., Pillay, N., Heinrichs, M. Schoepf, I., Schradin, C. 2015. Personality does not constrain social and behavioural flexibility in African striped mice. Behavioral Ecology and Sociobiology 69, 1237-1249.


45. Schoepf, I., Schradin, C. 2014. Arginine vasopressin plasma levels change seasonally in African striped mice but do not depend on reproductive tactics. General and Comparative Endocrinology 204, 43-48.


44. Schradin, C., Raynaud, J., Arrive, M. & Blanc, S. 2014. Leptin levels in free ranging striped mice (Rhabdomys pumilio) increase when food decreases. General and Comparative Endocrinology 206, 139-145.


43. Schradin, C., Larke, R. H. & Bales, K. L. 2014. Growing up in the family or growing up alone influences behavior and hormones, but not arginine vasopressin receptor 1a expression in male African striped mice. Physiology & Behavior, 129, 205-213.


42. Raynaud, J. & Schradin, C. 2014. Experimental increase of testosterone increase boldness and decrease anxiety in male African striped mouse helpers. Physiology & Behavior 129, 57-63.


41. Schradin, C. & Pillay, N. 2014. Absence of reproductive suppression in young adult female striped mice living in their natal family. Animal Behaviour, 90, 141-148.


40. Raynaud, J. & Schradin, C. 2013. Regulation of male prolactin levels in an opportunistically breeding species, the African striped mouse. Journal of Zoology, 290, 287-292.


39. Schoepf, I. & Schradin, C. 2013. Endocrinology of sociality: Comparisons between sociable and solitary individuals within the same population of African striped mice. Hormones and Behavior, 64, 89-94.


38. Schradin, C. 2013. Intra-specific variation in social organisation by genetic variation, developmental plasticity, social flexibility or only extrinsic factors. Philisophical Transactions of the Royal Society B.

37. Rymer, T., Pillay, N., Schradin, C., 2013. Extinction or Survival? Behavioral Flexibility in Response to Environmental Change in the African Striped Mouse Rhabdomys. Sustainability 5: 163-186.

36. Schradin, C., Kenkel, W., Krackow, S., Carter, C., 2013. Staying put or leaving home: endocrine, neuroendocrine and behavioral consequences in male African striped mice. Hormones and Behavior 63: 136-143.

35. Schoepf. I & Schradin C. 2012. Differences in social behaviour between group-living and solitary African striped mice, Rhabdomys pumilio. Animal Behaviour. 84: 1159-1167.

34. Raynaud, J., Müller, . & Schradin, C. 2012. Experimental increase of testosterone levels in free-ranging juvenile male African striped mice (Rhabdomys pumilio) induces physiological, morphological, and behavioral changes. General and Comparative Endocrinology 178: 108-115.

33. Schradin, C., Eder, S. & Müller, K. 2012. Differential investment into testes and sperm production in alternative male reproductive tactics of the African striped mouse (Rhabdomys pumilio). Hormones and Behavior 61: 686-695.

32. Schoepf, I. & Schradin, C. 2012. Better off alone! Reproductive competition and ecological constraints determine sociality in the African striped mouse (Rhabdomys pumilio). Journal of Animal Ecology 81: 649-656.

30. Schradin, C. & Yuen, C.H. 2011. Hormone levels of male African striped mice change as they switch between alternative reproductive tactics. Hormones and Behavior. 60: 676-680.

29. Schradin, C .& Lindholm, A.K. 2011. Relative fitness of alternative male reproductive tactics in a mammal varies between years. Journal of Animal Ecology 80, 908-917.

28. Solmsen, N., Johannesen, J. & Schradin C., 2011. Highly asymmetric finescale genetic structure between sexes of African striped mice indicate alternative male dispersal tactics Molecular Ecology 20: 1624-1634.

27. Schradin, C., Schneider, C. & Lindholm, A.K. 2010. The nasty neighbor in the striped mouse (Rhabdomys pumilio) steals paternity and elicits aggression. Frontiers in Zoology 7:19.

26. Scantlebury, M., Bennett, N., Pillay, N. & Schradin, C. 2010. Basking is affected by season and influences oxygen consumption in desert-living striped mice. Journal of Zoology. 281: 132-139.

25. Wilson, L.A.B., Schradin, C., Mitgutsch, C., Galliari, F.C., Andrea Mess, A., Sánchez-Villagra, M.R. accept. Skeletogenesis in the African striped mouse (Rhabdomys pumilio) and a study of sequence heterochrony in rodent evolution. Organisms Diversity and Evolution.

24. Schradin, C., König, B. & Pillay, N. 2010. Reproductive competition favours solitary living while ecological constraints impose group-living in African striped mice. Journal of Animal Ecology. 73: 515-521.

23. Schradin, C., Schmohl, G., Rödel, H. G., Schoepf, I., Treffler, S. M., Brenner, J., Bleeker, M., Schubert, M., König, B. & Pillay, N. Female home range size is regulated by resource distribution and intraspecific competition: a long-term field study. Animal Behaviour 79: 195-203.

22. Schradin, C., Schneider, C., Yuen, C.H. (2009). Age at puberty in male African striped mice: the impact of food, population density and the presence of the father. Functional Ecology 23:1004-1013.

21. Schradin, C., Kinahan, A.A., Pillay, N. (2009). Cooperative breeding in groups of synchroneously mating females and evolution of large testes to avoid sperm depletion in African striped mice. Biology of Reproduction 81: 111-117.

20. Schubert, M., Pillay, N. & Schradin, C. (2009). Parental and alloparental care in a polygynous mammal. Journal of Mammalogy90: 724-731.

19. Schradin, C. , Scantlebury, M., Pillay, N. & König, B. (2009). Testosterone levels in dominant sociable males are lower than in solitary roamers: physiological differences between three male reproductive tactics in a sociably flexible mammal. American Naturalist 173: 376-388.

18. Schradin, C. (2008). Seasonal changes in testosterone and corticosterone levels in four social categories of a desert dwelling sociable rodent. Hormones and Behavior 53: 573-579.

17. Schradin, C. (2008). "Differences in prolactin levels between three alternative male reproductive tactics in striped mice (Rhabdomys pumilio)." Proc R Soc B 275: 1047-1052.

16. Schradin, C. 2007. Comments to K.E. Wynne-Edwards & M.E. Timonin. Paternal care in rodents: Weakening support for hormonal regulation of the transition to behavioral fatherhood in rodent animal models of biparental care. Horm & Behav 52: 114-121. Hormones and Behavior 52: 557-559.

15. Schradin C, Krackow S, Schubert M, Keller C, Schradin B, Pillay N. 2007. Regulation of activity in desert-living striped mice: The importance of basking Ethology 113:606-614

14. Schradin C. 2006. Information transfer about food locations is not a benefit of group living in the solitary foraging striped mouse (Rhabdomys pumilio). J Ethol

13. Schradin C, 2006. Whole day follows of the striped mouse. J Ethol 24:37-43.

12. Schradin C, Pillay N, 2006. Female striped mice (Rhabdomys pumilio) change their home ranges in response to seasonal variation in food availability. Behav Eco. 17: 452-458.

11. Schradin C, Schubert M, Pillay N, 2006. Winter huddling groups in the striped mouse. Can J Zool. 84, 693-698.

10. Scantlebury M, Bennett NC, Speakman JR, Pillay N, Schradin C 2006. Huddling in groups leads to daily energy savings in free-living African four-striped grass mice, Rhabdomys pumilio. Funct Ecol 20:166-173

9. Schradin C, 2005. When to live alone and when to live in groups: ecological determinants of sociality in the African striped mouse (Rhabdomys pumilio, Sparrman, 1784). Belgian Journal of Zoolog 135: 77-82.

8. Schradin, C. 2005. Nest-site competition in two diurnal rodents from the Succulent Karoo of South Africa. Journal of Mammalogy 86: 757-762.

7. Schradin C, Pillay N, 2005. Demography of the striped mouse (Rhabdomys pumilio) in the succulent karoo. Mammalian Biology 70: 84-92.

6. Schradin C, Pillay N, 2005. The influence of the father on offspring development in the striped mouse. Behavioral Ecology. 16:450-455.

5. Schradin C, Pillay N, 2005. Intraspecific variation in the spatial and social organization of the African striped mouse. Journal of Mammalogy 86: 99-107.

4. Schradin C, 2004. Territorial defense in a group living solitary forager: who, where against whom? Behavioral Ecology and Sociobiology 55:439-446.

3. Schradin C, Pillay N, 2004a. Prolactin levels in paternal striped mouse (Rhabdomys pumilio) fathers. Physiology and Behavior 81:43-50.

2. Schradin C, Pillay N, 2004b. The striped mouse (Rhabdomys pumilio) from the succulent karoo of South Africa: A territorial group living solitary forager with communal breeding and helpers at the nest. Journal of Comparative Psychology 118:37-47.

1. Schradin C, Pillay N, 2003. Paternal care in the social and diurnal striped mouse (Rhabdomys pumilio): laboratory and field evidence. Journal of Comparative Psychology 117:317-324.


PhD theses

  • Schubert, M. 2009. The Social System of the Round-Eared Sengi (Macroscelides proboscideus), University of Bayreuth, Germany.

MSc theses


Die Biologie des Vaters

Popular science book in German, Filander Verlag Fürth, 29.90 Euro.